We follow Mahiro Oyama, an unemployed good-for-nothing shut-in, who not only hasn't left the house in two years, but is also exceedingly addicted to gross video games. One afternoon, he wakes up and notices that his body feels a little off. Finding that strange, he goes to reach for his tablet, but what does he see in his reflection, but an adorable young lady. I wonder what lies in store for the "former" big brother, Mahiro as he's thrust headfirst into his new life as a girl.

As Mahiro stumbles through his new life as a girl, he finds himself gradually beginning to get into the swing of things. Realizing this, however, prompts him to try and reclaim some of his masculinity. He tries, in futility, to engage his core and do a sit-up, but ends up with a stomachache instead. Seeing his condition worsen rapidly before her eyes, Mihari has an epiphany. She realizes that his pain may actually have something to do with fact that almost a month has passed since Mahiro turned into a girl...

In a rare turn of events, Mihari wakes up one morning to find Mahiro has gotten up before her. Since the corner store was going to start selling clear files from his favorite anime that day, he had woken up early to go pick them up before they sold out. Mihari is overwhelmed with joy to see Mahiro going outside of his own accord. On his way back from buying the clear files, Mahiro stops by an arcade for the first time since middle school. He's enjoying a rhythm game when suddenly he senses a man standing watching him from behind. A few minutes go by when he feels a tap on the shoulder...

It's finally winter break and Mahiro is excited to get into his Winter Vacation Mode. Unbeknownst to him, however, Mihari invites Momiji and the others over for a sleepover. She hopes this will prevent a potential relapse into his old, shut-in ways.